Observations by royh

Pileus 4-6 cm broad, convex to plane, dry, finely subtomentose to barely subvelutinous, dark brown to cocoa brown (6F7-6, 6E7-6), becoming areolate with age. NH4 - a blue green flash then light lavender. Flesh white, staining pale blue then grayish fuscous to dark gray. Lamellae decurrent, subdistant, sometimes with anastomoses, up to 1 cm broad, bright yellow (2A7-6, 3A7-6), staining blue green when bruised, with even to uneven edges. Stipe 4-5 cm long, 8-15 mm broad, equal, usually curved, dry, white, subpubescent to matted subtomentose, slowly brownish gray from handling, with interior white, solid, staining pale blue green then dark grayish fuscous; basal mycelium white. Determined by: M.-A. Neves
GenBank: JQ003684 (nrLSU); JQ003621 (ITS)

Dark gray cap and stipe. Stipe white at base with reticulum above. Tubes dull yellow, staining red. GenBank RPB2 = MH712034; ITS = MK248715
Light micrographs: Spores, pileipellis, stipitipellis, cheilocystidia

Pileus 2-8 cm broad, convex to plano-convex, dry, subtomentose to matted then finely areolate with age, black to gray becoming brown to cocoa brown (6E6). Flesh white, slowly dull pinkish tan with exposure, with mild odor and taste. Tubes adnexed, white when young, light grayish yellow (4B4) then darker grayish yellow (4C4), slowly staining dingy reddish with bruising, with pores likewise, deep reddish bluish with NH4OH. Stipe 4-7 cm long, 5-10 mm broad, equal to subclavate and then tapered at base, dry, black, reticulate nearly to base, dark grayish to brownish near base, white at base, with interior white, slowly pale to dull pinkish tan. Spore print reddish brown. Spores violet bluish in KOH.
NB: One basidiome with finely scissurate scabrous stipe, otherwise similar to those with reticulate stipe (9784b). GenBank RB2 = MH712036

Pileus 8-9 cm broad, plano-convex, dry, finely appressed squamulose scaly, deep red with some olive to yellow in areas near margin, with a slight projecting sterile margin. Flesh very pale yellow to yellowish white, slowly cyanescent, with mild odor and taste. Tubes adnexed, olive brown, cyanescent, with pores pale orangish red and cyanescent. Stipe 6 cm long, 2 cm broad, with a slight submarginate bulb, yellowish white with some pale greenish overtones and finely striate, lightly cyanescent, with an obvious marginate limb at edge of bulb, appressed to suberect squamulose scaly below limb, with scales a deep red to brown, with egg yolk yellow (4B8) tomentum at extreme base (attachment point), with interior yellowish white deeply cyanescent but slowly.

This is a button stage of 8746, 8776 showing the universal veil attachment. finely squamulose on stipe base and pileus, staining blue. Bright yellow tubes, hardly bluing if at all. 3 cm broad, 4.5 cm tall.

Pileus 6.5-9 cm broad, plano-convex, dry, deep brownish red on disc, dirty yellowish at margin, overlain with agglutinated and appressed dark brown squamules, staining blue when bruised. Flesh pale lemon yellow, slowly blue when exposed, 2 cm thick. Tubes adnexed, bright yellow, bluing, with pores olive yellow (3C8), bruising blue, 2-3/mm. Stipe 9.5 cm long, 1.5-3 cm broad, slightly flattened, equal to subclavate with a pinched base, bright lemon yellow to chrome yellow and fibrillose streaked above marginate limb, staining blue, with reddish to reddish brown agglutinated and appressed squamules below marginate limb, overlaying a pale yellowish ground, staining blue, with interior pale lemon yellow, staining blue, solid, with yellow basal mycelium.

Pileus 7-9 cm broad, convex to plano-convex, dry, finely appressed squamulose, with squamules brown to reddish brown, overlying a dull red disc, dull yellow to bright yellow at margin, staining blue, with even margin. Flesh pale yellow to pale lemon yellow, 1-2 cm thick, staining blue when exposed, with mild odor and slowly unpleasant to nearly bitter taste. Hymenophore adnexed to depressed around stipe, bright yellow to bright greenish yellow (2A-B7,6), staining blue-green when bruised at first, then pores becoming brown. Stipe 6.5-9.5 cm long, 2-2.7 cm broad, equal to subclavate to clavate with a submarginate bulb clearly defined and tapering to base below marginate rim; surface bright lemon yellow above and heavily pruinose to subfloccose, matted toward base, pale pinkish red with fine appressed brown squamules below marginate rim (as in pileus); interior solid, yellow, staining blue.

Pileus whitish, becoming dull yellowish or pinkish with age, with suede-like texture and tacky-sticky at first, somewhat subviscid with age. White lacunose stipe, all parts unchanging. Tubes whitish at first, pinkish vinaceous with age. Quite similar to material from Fraser Island.

Pileus with suede-like texture when young, nearly pure white with some pale pinkish tones where pressed, becoming pale yellow and matted. Stipe pure white and alveolate-reticulate, developing yellow tones near base with age.

Pileus 4-8 cm broad, convex to plano-convex to plane, moist and sticky but not glutinous or with slime, with suede-like texture at first, this soon matted and disappearing, white to a pale yellow (4A3) at first, soon with more yellow and some pale pinkish tints (7A-B5,4), with sterile appendiculate marginal tissue when young. Flesh white, unchanging, with mild odor and bitter taste. Tubes adnexed, whitish at first, soon porphyry pinkish, bruising pale brownish. Stipe 5-12 cm long, 1-1.5 cm broad, strict of curved, equal to slightly broader below, often tapered at base, white and lacunose reticulate at first, soon with yellow or orangish yellow stains, moist and sticky with some gluten here and there, with interior white, unchanging, solid to pithy becoming hollow.

Pileus 3-6 cm broad, convex to plano-convex, moist and sticky but not with gluten, with suede-like texture, white to very pale yellow, sometimes with pale brown to pinkish brown areas, with white, sterile, appendiculate margin fragments. Odor mild to slightly fragrant, with bitter taste from sticky/subviscid areas. Flesh white, unchanging. Tubes adnexed, white at first, soon becoming porphyry pinkish. Stipe 4-9 cm long, 7-13 mm broad, with some gluten above, soon disappearing, white overall at first, lacunose-reticulate, developing scattered yellowish stains toward and at base with age, with interior becoming hollow, white, with yellowish tints in base with age, with white basal mycelium.

Pileus 1-5 cm broad, convex, viscid (wet weather), or glutinous when very young, soon smooth leather-like-areolate (nearly like suede, but not cracking), pale yellow (4A4) on disc, whiter toward margin, with white appendiculate veil remnants at margin. Flesh white, unchanging with mild odor and bitter taste. Tubes adnexed, whitish or with slight pinkish tint (young). Stipe 5-9 cm long, 8-10 mm broad, subclavate, white, lacunose reticulate, with reticulum becoming gelatinous from handling, developing orangish yellow stains with age, with interior solid, white, unchanging.

Pileus 6.5-9 cm broad, plano-convex to plane, dry, matted subvelutinous to matted subtomentose, red brown, with even margin. Flesh grayish vinaceous to lilac gray, unchanging. Tubes adnexed, rusty brown to pinkish brown, unchanging, with pores rusty brown to pinkish brown and unchanging. Stipe 3-5 cm long, 1-8-2 cm broad, equal, dry scissurate scaly, brownish lilac to grayish lilac to light rusty brown over a silvery background, unchanging, with interior lilac gray, unchanging, white at the base.

Pileus 3-9 cm broad, convex to plane, dry, dark brown (7,8F8,7) to nearly black in some, becoming brown (7E6,5), even at margin. Flesh white to pale lilac with pinkish brown to brownish lilac marbling/mottling, with mild odor and mild to slightly unpleasant slowly slightly bitterish taste. Tubes adnexed, lilac whitish when young, soon grayish orange (6B3) to light brown (6D4), with pores violet brown (11F5) when young, becoming brown (7E6,5) to cocoa brown (6E7) with age. Stipe 2.5-6 cm long, 1-2 cm broad, equal, strict or curved, dry, finely scabrous on a pale lilac ground, with scabers a dull brown, with interior whitish to grayish lilac and mottled, becoming streaked with pale brown, apparently with white basal mycelium but this obscured by sand.

Pileus 7-8 cm broad, convex to plano-convex, dry to viscid (wet weather), very finely matted, dark brown (9F8) to lilac brown (11D4) (especially toward margin). Flesh mottled sordid lilac with white, unchanging, with mild odor and taste. Tubes adnexed to deeply depressed, mature and light cinnamon brown, staining a darker brown. Stipe 4-6 cm long, 1.3-1.8 cm broad, equal, dry, finely scabrous, pale lilac with pinkish brown to pinkish lilac scabers, darker below, with interior mottled pale lilac in areas with light brown, with some apparent light brownish orange staining. Apparently whitish at base, but with too much sand to determine color.

Brown staining 'cyanescens' entity. Pileus heavily matted fibrillose; stipe likewise with annular zone.
Davoodian, N. et al. 2018. A global view of Gyroporus: molecular phylogenetics, diversity patterns, and new species. Mycologia. 110 (/doi.org/10.1080/00275514.2018.1511339)

Pileus 1.5-5 cm broad, convex to plano-convex, dry, matted tomentose to matted fibrillose, grayish yellow (4B-C5,4), even at margin. Flesh white, slowly pale brown when exposed, with mild odor and taste. Tubes adnate to adnexed, pale yellowish white to very pale yellow, with pores bruising brown (not tubes). Stipe 1.5-3 cm long, 9-15 mm broad, dry, matted fibrillose, with velar zone, grayish yellow (as in pileus), bruising pale brown, with interior white, becoming brown when exposed, soon hollow and/or chambered.
Davoodian, N. et al. 2018. A global view of Gyroporus: molecular phylogenetics, diversity patterns, and new species. Mycologia. 110 (/doi.org/10.1080/00275514.2018.1511339)

Pileus 9.2-9.5 cm broad, plano-convex, even, dry to slightly tacky, tomentose to matted tomentose to nearly subtomentose, orange brown (6C7-6-5), with a sterile irregular appendiculate margin, whitish in color or with some yellowish pigments as in disc. Flesh white, unchanging when exposed, 2 cm thick, with mild odor or no odor and mild taste. Tubes deeply adnexed to nearly free, pinkish flesh color, staining brownish; pores 1 mm diam. Stipe 8 cm long, 1.7 cm broad, equal, tapering at base, dry, coarsely alveolate-reticulate, white above, with some pale yellow toward base, staining red at base from handling, with interior white, soon chambered by insect larvae, apparently staining yellow in cortex after exposure; basal mycelium covered with sand.
PARATYPE
GenBank: ITS (KP242166)

Pileus 3.5-4.5 cm broad when expanded, glutinous to viscid, becoming minutely areolate tomentose as gluten dissipates; margin with white appendiculate veil remnants; colored 7E8 at first then 7D7. Flesh white, unchanging, up to 1 cm deep. Tubes white when young, soon pinkish brown (8D4), unchanging, depressed to deeply depressed around stipe; pores concolorous, not staining, up to 1 mm to 1.5 mm broad. Stipe 4.5-8 cm x 6-10 mm, equal, strict or curved, white with alveolate reticulate ridges, the latter white above, viscid below and brownish from gluten and handling, white at base with white rhizomorphs. Odor mild, taste mild. Spores very finely & uniformly rugulose to irregularly granular.
PARATYPE
GenBank: nr28s (KP242247)

Pileus 3.5-7.5 cm broad, convex to obtusely conico-convex, dry, with fine erect pyramidal squamules to tomentose, brown to cocoa brown (6E8,7), with sterile appendiculate margin when young. Flesh white, unchanging with mild odor and taste. Tubes adnexed to deeply adnexed, very pale pinkish at first, becoming vinaceous pink with maturity, staining slightly a brownish pink. Stipe 3.5-6 cm long, 1-1.5 cm broad, strict or slightly curved, equal or equal and tapered at base, lacunose-reticulate, dry, with brown reticulum below and yellowish between 'nets', whitish to pale yellow above veil rupture line when young, the 'nets' becoming more pinkish brown with age, otherwise pale yellow near apex, with interior white, unchanging, pithy, very pale yellow at base, with white basal mycelium. PARATYPE
GenBank: nr28s (MT921383); rpb1 (MT932084); rpb2 (MT928122)

Pileus 4.5 cm broad, plano-convex, dry, deep dark red, subvelutinous, mottled slightly with small dark yellow spots. Flesh pale yellow, cyanescent then obscurely white, with mild odor and taste. Tubes adnexed, bright yellow to dark yellow, cyanescent with pores cyanescent. Stipe 4 x 7 cm, curved, slightly broader below, dry, uniformly deep red with a white base, heavily pruinose above, fibrillose striate below, cyanescent, with interior yellow, cyanescent, deep reddish around larval tunnels.

Pileus 3.5-4 cm broad, convex, dry, subtomentose, very finely areolate, deep red. Flesh pale yellow, rapidly cyanescent, with mild odor and taste. Tubes adnexed, greenish yellow, cyanescent, pore likewise. Stipe 4-5 cm long, 6-9 mm broad, equal and slightly expanded at base and then tapering to a blunt point, yellow at extreme apex, then red overall and subpruinose, white at base, with interior yellow cyanescent, solid. Vaguely reminiscent of B. obscurecoccineus, but is cyanescent.
This is the TYPE

Pileus with white, flattened cottony patches with pink tints between, slightly sticky, 3-4 cm broad, plano-convex. Tubes adnexed, pinkish vinaceous. Stipe 6-7 cm long, 5-7 mm broad, equal to slightly broader below, with delicate white sublacunose reticulum, with some yellow tones in flesh at stipe base.